*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
    --primary: #181d26;
    --primary-active: #0d1218;
    --canvas: #ffffff;
    --surface-soft: #f8fafc;
    --surface-strong: #e0e2e6;
    --surface-dark: #181d26;
    --hairline: #dddddd;
    --ink: #181d26;
    --body: #333840;
    --muted: #41454d;
    --on-primary: #ffffff;
    --link: #1b61c9;
    --link-active: #1a3866;
    --signature-coral: #aa2d00;
    --signature-forest: #0a2e0e;
    --signature-cream: #f5e9d4;
    --signature-peach: #fcab79;
    --signature-mint: #a8d8c4;
    --signature-yellow: #f4d35e;
    --rounded-xs: 2px;
    --rounded-sm: 6px;
    --rounded-md: 10px;
    --rounded-lg: 12px;
    --spacing-xs: 8px;
    --spacing-sm: 12px;
    --spacing-md: 16px;
    --spacing-lg: 24px;
    --spacing-xl: 32px;
    --spacing-xxl: 48px;
    --spacing-section: 96px;
    --max-width: 1280px;
}

html { font-size: 16px; scroll-behavior: smooth; }

body {
    font-family: 'Inter', -apple-system, BlinkMacSystemFont, 'Segoe UI', Roboto, Oxygen, Ubuntu, Cantarell, 'Fira Sans', 'Droid Sans', 'Helvetica Neue', sans-serif;
    font-size: 14px;
    font-weight: 400;
    line-height: 1.25;
    color: var(--body);
    background: var(--canvas);
}

a { color: var(--link); text-decoration: none; }
a:hover { color: var(--link-active); }

img { max-width: 100%; display: block; }

.container {
    max-width: var(--max-width);
    margin: 0 auto;
    padding: 0 var(--spacing-xxl);
}

@import url('https://fonts.googleapis.com/css2?family=Inter:ital,opsz,wght@0,14..32,400;0,14..32,500;0,14..32,600;1,14..32,400&display=swap');

.top-nav {
    position: sticky;
    top: 0;
    z-index: 100;
    height: 64px;
    background: var(--canvas);
    border-bottom: 1px solid var(--hairline);
}

.nav-inner {
    max-width: var(--max-width);
    margin: 0 auto;
    padding: 0 var(--spacing-xxl);
    height: 100%;
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: var(--spacing-lg);
}

.nav-logo {
    font-size: 18px;
    font-weight: 600;
    color: var(--ink);
    letter-spacing: -0.3px;
    flex-shrink: 0;
}

.nav-logo:hover { color: var(--ink); }

.nav-links {
    display: flex;
    list-style: none;
    gap: var(--spacing-lg);
    margin: 0;
}

.nav-links a {
    font-size: 14px;
    font-weight: 400;
    color: var(--body);
    transition: color 0.15s;
}

.nav-links a:hover,
.nav-links a.nav-active { color: var(--ink); font-weight: 500; }

.nav-hamburger {
    display: none;
    flex-direction: column;
    gap: 5px;
    background: none;
    border: none;
    cursor: pointer;
    padding: 4px;
}

.nav-hamburger span {
    display: block;
    width: 22px;
    height: 2px;
    background: var(--ink);
    border-radius: 2px;
    transition: all 0.2s;
}

.hero-band {
    padding: var(--spacing-section) 0;
    background: var(--canvas);
}

.hero-band .container { max-width: 860px; }

.hero-eyebrow {
    font-size: 13px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: var(--muted);
    text-transform: uppercase;
    margin-bottom: var(--spacing-md);
}

.hero-band h1 {
    font-size: 40px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    margin-bottom: var(--spacing-lg);
}

.hero-band p {
    font-size: 18px;
    font-weight: 400;
    line-height: 1.6;
    color: var(--body);
    max-width: 640px;
    margin-bottom: var(--spacing-xxl);
}

.btn-primary {
    display: inline-flex;
    align-items: center;
    gap: var(--spacing-xs);
    background: var(--primary);
    color: var(--on-primary);
    font-size: 16px;
    font-weight: 500;
    line-height: 1.4;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: none;
    cursor: pointer;
    transition: background 0.15s;
    text-decoration: none;
}

.btn-primary:hover { background: var(--primary-active); color: var(--on-primary); }

.btn-secondary {
    display: inline-flex;
    align-items: center;
    gap: var(--spacing-xs);
    background: var(--canvas);
    color: var(--ink);
    font-size: 16px;
    font-weight: 500;
    line-height: 1.4;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: 1px solid var(--hairline);
    cursor: pointer;
    transition: border-color 0.15s;
    text-decoration: none;
}

.btn-secondary:hover { border-color: var(--ink); color: var(--ink); }

.btn-row { display: flex; gap: var(--spacing-md); flex-wrap: wrap; align-items: center; }

.section-band {
    padding: var(--spacing-section) 0;
}

.section-band--soft { background: var(--surface-soft); }
.section-band--dark { background: var(--surface-dark); }
.section-band--strong { background: var(--surface-strong); }

.section-label {
    font-size: 13px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: var(--muted);
    text-transform: uppercase;
    margin-bottom: var(--spacing-md);
}

.section-label--light { color: rgba(255,255,255,0.6); }

.section-heading {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    margin-bottom: var(--spacing-lg);
}

.section-heading--light { color: var(--on-primary); }

.section-sub {
    font-size: 16px;
    line-height: 1.6;
    color: var(--body);
    max-width: 600px;
    margin-bottom: var(--spacing-xxl);
}

.section-sub--light { color: rgba(255,255,255,0.75); }

.cards-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
}

.article-card {
    background: var(--canvas);
    border-radius: var(--rounded-md);
    border: 1px solid var(--hairline);
    overflow: hidden;
    transition: box-shadow 0.2s;
}

.article-card:hover { box-shadow: 0 4px 16px rgba(24,29,38,0.08); }

.article-card__img {
    aspect-ratio: 16/9;
    overflow: hidden;
}

.article-card__img img {
    width: 100%;
    height: 100%;
    object-fit: cover;
}

.article-card__body {
    padding: var(--spacing-md);
}

.article-card__tag {
    display: inline-block;
    font-size: 12px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: var(--muted);
    text-transform: uppercase;
    margin-bottom: var(--spacing-xs);
}

.article-card__title {
    font-size: 18px;
    font-weight: 500;
    line-height: 1.4;
    color: var(--ink);
    margin-bottom: var(--spacing-xs);
}

.article-card__title a { color: var(--ink); }
.article-card__title a:hover { color: var(--link); }

.article-card__meta {
    font-size: 13px;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.article-card__excerpt {
    font-size: 14px;
    line-height: 1.6;
    color: var(--body);
}

.signature-coral-card {
    background: var(--signature-coral);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-xxl);
}

.signature-forest-card {
    background: var(--signature-forest);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-xxl);
}

.signature-card__eyebrow {
    font-size: 13px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: rgba(255,255,255,0.65);
    text-transform: uppercase;
    margin-bottom: var(--spacing-md);
}

.signature-card__title {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--on-primary);
    margin-bottom: var(--spacing-md);
}

.signature-card__text {
    font-size: 14px;
    line-height: 1.6;
    color: rgba(255,255,255,0.8);
    margin-bottom: var(--spacing-xxl);
    max-width: 520px;
}

.signature-cards-grid {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: var(--spacing-lg);
}

.species-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
}

.species-card {
    background: var(--canvas);
    border-radius: var(--rounded-md);
    border: 1px solid var(--hairline);
    overflow: hidden;
}

.species-card__img {
    aspect-ratio: 4/3;
    overflow: hidden;
}

.species-card__img img {
    width: 100%;
    height: 100%;
    object-fit: cover;
}

.species-card__body { padding: var(--spacing-md); }

.species-card__genus {
    font-size: 12px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: var(--muted);
    text-transform: uppercase;
    margin-bottom: 4px;
}

.species-card__name {
    font-size: 16px;
    font-weight: 500;
    line-height: 1.4;
    color: var(--ink);
    font-style: italic;
    margin-bottom: var(--spacing-xs);
}

.species-card__common {
    font-size: 13px;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.species-card__desc {
    font-size: 13px;
    line-height: 1.6;
    color: var(--body);
}

.status-badge {
    display: inline-block;
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.12px;
    padding: 3px 8px;
    border-radius: var(--rounded-sm);
    margin-top: var(--spacing-xs);
}

.status-badge--lc { background: #d1f5d3; color: #006400; }
.status-badge--nt { background: #fff3cd; color: #856404; }
.status-badge--vu { background: #fde8cc; color: #7a3a00; }
.status-badge--en { background: #f8d7da; color: #842029; }

.cream-band {
    background: var(--signature-cream);
    padding: var(--spacing-section) 0;
}

.cream-band__inner {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: var(--spacing-xxl);
    align-items: center;
}

.cream-band__title {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    margin-bottom: var(--spacing-lg);
}

.cream-band__text {
    font-size: 14px;
    line-height: 1.7;
    color: var(--body);
    margin-bottom: var(--spacing-lg);
}

.phenology-table {
    width: 100%;
    border-collapse: collapse;
    font-size: 13px;
}

.phenology-table th {
    font-weight: 500;
    color: var(--muted);
    text-align: left;
    padding: 8px 12px;
    border-bottom: 1px solid var(--hairline);
    font-size: 12px;
    letter-spacing: 0.1px;
}

.phenology-table td {
    padding: 10px 12px;
    border-bottom: 1px solid var(--hairline);
    color: var(--body);
}

.phenology-table tr:last-child td { border-bottom: none; }

.month-bar {
    display: flex;
    gap: 2px;
}

.month-dot {
    width: 14px;
    height: 14px;
    border-radius: 3px;
    background: var(--hairline);
}

.month-dot--active { background: var(--signature-coral); }

.contact-section {
    padding: var(--spacing-section) 0;
    background: var(--canvas);
}

.contact-grid {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: var(--spacing-xxl);
}

.contact-form {
    display: flex;
    flex-direction: column;
    gap: var(--spacing-md);
}

.form-group {
    display: flex;
    flex-direction: column;
    gap: 6px;
}

.form-group label {
    font-size: 13px;
    font-weight: 500;
    color: var(--ink);
}

.form-group input,
.form-group textarea {
    background: var(--canvas);
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-sm);
    padding: 12px 16px;
    font-size: 14px;
    font-family: inherit;
    color: var(--ink);
    height: 44px;
    transition: border-color 0.15s;
}

.form-group textarea { height: auto; min-height: 120px; resize: vertical; }

.form-group input:focus,
.form-group textarea:focus {
    outline: none;
    border-color: #458fff;
}

.btn-submit {
    display: inline-flex;
    align-items: center;
    background: var(--primary);
    color: var(--on-primary);
    font-size: 16px;
    font-weight: 500;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: none;
    cursor: pointer;
    transition: background 0.15s;
    align-self: flex-start;
}

.btn-submit:hover { background: var(--primary-active); }

.contact-info h3 {
    font-size: 24px;
    font-weight: 400;
    color: var(--ink);
    margin-bottom: var(--spacing-lg);
}

.contact-info p {
    font-size: 14px;
    line-height: 1.7;
    color: var(--body);
    margin-bottom: var(--spacing-md);
}

.contact-detail {
    display: flex;
    flex-direction: column;
    gap: var(--spacing-sm);
    margin-top: var(--spacing-lg);
}

.contact-detail span {
    font-size: 14px;
    color: var(--body);
    display: flex;
    gap: var(--spacing-xs);
}

.cta-band {
    background: var(--surface-strong);
    padding: var(--spacing-section) 0;
}

.cta-band__inner {
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: var(--spacing-xxl);
}

.cta-band__title {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    max-width: 480px;
}

.footer {
    background: var(--canvas);
    border-top: 1px solid var(--hairline);
    padding: var(--spacing-section) 0 var(--spacing-xl);
}

.footer-inner {
    max-width: var(--max-width);
    margin: 0 auto;
    padding: 0 var(--spacing-xxl);
}

.footer-cols {
    display: grid;
    grid-template-columns: 2fr 1fr 1fr 1fr;
    gap: var(--spacing-xl);
    margin-bottom: var(--spacing-xxl);
}

.footer-brand {
    font-size: 16px;
    font-weight: 600;
    color: var(--ink);
    display: block;
    margin-bottom: var(--spacing-sm);
}

.footer-desc {
    font-size: 13px;
    line-height: 1.6;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.footer-contact {
    font-size: 13px;
    line-height: 1.8;
    color: var(--muted);
}

.footer-contact a { color: var(--muted); }
.footer-contact a:hover { color: var(--link); }

.footer-heading {
    font-size: 13px;
    font-weight: 500;
    color: var(--ink);
    display: block;
    margin-bottom: var(--spacing-sm);
}

.footer-col ul { list-style: none; display: flex; flex-direction: column; gap: 6px; }
.footer-col ul a { font-size: 13px; color: var(--muted); }
.footer-col ul a:hover { color: var(--link); }

.footer-legal {
    display: flex;
    justify-content: space-between;
    align-items: center;
    padding-top: var(--spacing-xl);
    border-top: 1px solid var(--hairline);
    font-size: 13px;
    color: var(--muted);
}

.article-hero {
    padding: var(--spacing-section) 0 var(--spacing-xxl);
    background: var(--canvas);
}

.article-hero__tag {
    font-size: 12px;
    font-weight: 500;
    letter-spacing: 0.16px;
    color: var(--muted);
    text-transform: uppercase;
    margin-bottom: var(--spacing-md);
}

.article-hero h1 {
    font-size: 40px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    margin-bottom: var(--spacing-lg);
    max-width: 760px;
}

.article-hero__meta {
    font-size: 13px;
    color: var(--muted);
    margin-bottom: var(--spacing-xxl);
}

.article-hero__img {
    width: 100%;
    max-height: 480px;
    overflow: hidden;
    border-radius: var(--rounded-md);
}

.article-hero__img img { width: 100%; height: 100%; object-fit: cover; }

.article-body {
    padding: var(--spacing-section) 0;
}

.article-layout {
    display: grid;
    grid-template-columns: 1fr 280px;
    gap: var(--spacing-xxl);
    align-items: start;
}

.article-content h2 {
    font-size: 28px;
    font-weight: 400;
    line-height: 1.3;
    color: var(--ink);
    margin-top: var(--spacing-xxl);
    margin-bottom: var(--spacing-lg);
}

.article-content h3 {
    font-size: 20px;
    font-weight: 500;
    line-height: 1.4;
    color: var(--ink);
    margin-top: var(--spacing-xl);
    margin-bottom: var(--spacing-md);
}

.article-content p {
    font-size: 15px;
    line-height: 1.75;
    color: var(--body);
    margin-bottom: var(--spacing-lg);
}

.article-content ul, .article-content ol {
    padding-left: 24px;
    margin-bottom: var(--spacing-lg);
}

.article-content li {
    font-size: 15px;
    line-height: 1.75;
    color: var(--body);
    margin-bottom: 6px;
}

.article-content figure {
    margin: var(--spacing-xxl) 0;
}

.article-content figure img {
    width: 100%;
    border-radius: var(--rounded-md);
}

.article-content figcaption {
    font-size: 12px;
    color: var(--muted);
    margin-top: var(--spacing-xs);
    font-style: italic;
}

.article-content a { color: var(--link); }
.article-content a:hover { color: var(--link-active); }

.article-sidebar {
    position: sticky;
    top: 80px;
}

.sidebar-box {
    background: var(--surface-soft);
    border-radius: var(--rounded-md);
    padding: var(--spacing-md);
    margin-bottom: var(--spacing-lg);
    border: 1px solid var(--hairline);
}

.sidebar-box h4 {
    font-size: 13px;
    font-weight: 500;
    color: var(--ink);
    margin-bottom: var(--spacing-sm);
    text-transform: uppercase;
    letter-spacing: 0.1px;
}

.sidebar-box ul { list-style: none; display: flex; flex-direction: column; gap: 8px; }
.sidebar-box ul a { font-size: 13px; color: var(--body); }
.sidebar-box ul a:hover { color: var(--link); }

.page-hero {
    padding: var(--spacing-section) 0 var(--spacing-xxl);
    background: var(--canvas);
}

.page-hero h1 {
    font-size: 40px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--ink);
    margin-bottom: var(--spacing-md);
}

.page-hero p {
    font-size: 16px;
    line-height: 1.6;
    color: var(--body);
    max-width: 600px;
}

.page-body {
    padding: var(--spacing-section) 0;
}

.page-content h2 {
    font-size: 24px;
    font-weight: 400;
    color: var(--ink);
    margin-top: var(--spacing-xxl);
    margin-bottom: var(--spacing-lg);
}

.page-content h3 {
    font-size: 18px;
    font-weight: 500;
    color: var(--ink);
    margin-top: var(--spacing-xl);
    margin-bottom: var(--spacing-md);
}

.page-content p {
    font-size: 15px;
    line-height: 1.75;
    color: var(--body);
    margin-bottom: var(--spacing-md);
}

.page-content ul, .page-content ol {
    padding-left: 24px;
    margin-bottom: var(--spacing-md);
}

.page-content li {
    font-size: 15px;
    line-height: 1.75;
    color: var(--body);
    margin-bottom: 4px;
}

.page-content a { color: var(--link); }

.cookie-banner {
    position: fixed;
    bottom: 0;
    left: 0;
    right: 0;
    z-index: 200;
    background: var(--surface-dark);
    color: var(--on-primary);
    padding: var(--spacing-md) var(--spacing-xxl);
    display: none;
    align-items: center;
    justify-content: space-between;
    gap: var(--spacing-lg);
    flex-wrap: wrap;
}

.cookie-banner.is-visible { display: flex; }

.cookie-banner p {
    font-size: 13px;
    line-height: 1.5;
    color: rgba(255,255,255,0.8);
    flex: 1;
    min-width: 240px;
}

.cookie-banner a { color: rgba(255,255,255,0.6); text-decoration: underline; }

.cookie-btns { display: flex; gap: var(--spacing-xs); flex-shrink: 0; }

.btn-legal {
    background: var(--link);
    color: var(--on-primary);
    font-size: 13.12px;
    font-weight: 600;
    padding: 12px 10px;
    border-radius: var(--rounded-xs);
    border: none;
    cursor: pointer;
    white-space: nowrap;
}

.btn-legal--reject {
    background: transparent;
    color: rgba(255,255,255,0.7);
    border: 1px solid rgba(255,255,255,0.3);
}

.disclaimer {
    background: var(--surface-soft);
    border-left: 3px solid var(--hairline);
    padding: var(--spacing-md);
    border-radius: 0 var(--rounded-sm) var(--rounded-sm) 0;
    margin-bottom: var(--spacing-lg);
}

.disclaimer p {
    font-size: 13px;
    line-height: 1.6;
    color: var(--muted);
    margin: 0;
}

@media (max-width: 1024px) {
    .cards-grid { grid-template-columns: repeat(2, 1fr); }
    .species-grid { grid-template-columns: repeat(2, 1fr); }
    .footer-cols { grid-template-columns: 1fr 1fr; }
    .article-layout { grid-template-columns: 1fr; }
    .article-sidebar { position: static; }
    .cream-band__inner { grid-template-columns: 1fr; }
    .contact-grid { grid-template-columns: 1fr; }
    .cta-band__inner { flex-direction: column; align-items: flex-start; }
    .signature-cards-grid { grid-template-columns: 1fr; }
}

@media (max-width: 768px) {
    .container { padding: 0 var(--spacing-md); }
    .nav-inner { padding: 0 var(--spacing-md); }
    .nav-links { display: none; flex-direction: column; position: fixed; top: 64px; left: 0; right: 0; background: var(--canvas); border-bottom: 1px solid var(--hairline); padding: var(--spacing-lg); gap: var(--spacing-md); z-index: 99; }
    .nav-links.is-open { display: flex; }
    .nav-hamburger { display: flex; }
    .hero-band h1 { font-size: 28px; }
    .section-heading { font-size: 24px; }
    .cards-grid { grid-template-columns: 1fr; }
    .species-grid { grid-template-columns: 1fr; }
    .footer-cols { grid-template-columns: 1fr; }
    .footer-legal { flex-direction: column; gap: var(--spacing-xs); text-align: center; }
    .article-hero h1 { font-size: 28px; }
    .page-hero h1 { font-size: 28px; }
    .signature-card__title { font-size: 24px; }
    .cookie-banner { padding: var(--spacing-md); }
    .footer-inner { padding: 0 var(--spacing-md); }
    .signature-cards-grid { grid-template-columns: 1fr; }
    .hero-band { padding: var(--spacing-xxl) 0; }
    .section-band { padding: var(--spacing-xxl) 0; }
    .cream-band { padding: var(--spacing-xxl) 0; }
    .cta-band { padding: var(--spacing-xxl) 0; }
    .contact-section { padding: var(--spacing-xxl) 0; }
    .article-body { padding: var(--spacing-xxl) 0; }
    .page-body { padding: var(--spacing-xxl) 0; }
    .article-hero { padding: var(--spacing-xxl) 0 var(--spacing-xl); }
    .page-hero { padding: var(--spacing-xxl) 0 var(--spacing-xl); }
}